What other bird are Penguins closest relative ?

Answer 1

Biologists are actually somewhat divided on this topic. Some think the extinct Plotopteridae were their closest relatives. Among living species, it may be "higher waterbirds," which include storks, rails, and the seabirds. But none are really close relations. Grebes, conversely, are sometimes thought to be closely related to penguins, but actually are probably not.

Answer 2

Penguin Relatives

Answer 3

penguin,

originally the common name for the now extinct great auk of the N Atlantic and now used (since the 19th cent.) for the unrelated antarctic diving birds. Penguins, which are related most closely to the albatrosses, are the most highly specialized of all birds for marine life.

Answer 4

Modern penguins consititute two undisputed clades and another two more basal genera with more ambiguous relationships (Bertelli & Giannini 2005). The origin of the Spheniscinae lies probably in the latest Paleogene, and geographically it must have been much the same as the general area in which the order evolved: the oceans between the Australia-New Zealand region and the Antarctic (Baker et al. 2006). Presumedly diverging from other penguins around 40 mya (Baker et al. 2006), it seems that the Spheniscinae were for quite some time limited to their ancestral area, as the well-researched deposits of the Antarctic Peninsula and Patagonia have not yielded Paleogene fossils of the subfamily. Also, the earliest spheniscine lineages are those with the most southern distribution.

The genus Aptenodytes appears to be the basalmost divergence among living penguins; they have bright yellow-orange neck, breast, and bill patches, incubate by placing their eggs on their feet, and when they hatch, they are almost naked. This genus has a distribution centered on the Antarctic coasts and barely extends to some subantarctic islands today.

Pygoscelis contains species with a fairly simple black-and-white head pattern; their distribution is intermediate, centered on Antarctic coasts but extending somewhat northwards from there. In external morphology, these apparently still resemble the common ancestor of the Spheniscinae, as Aptenodytes' autapomorphies are in most cases fairly pronounced adaptations related to that genus' extreme habitat conditions. As the former genus, Pygoscelis seems to have diverged during the Bartonian, but the range expansion and radiation which lead to the present-day diversity probably did not occur until much later, around the Burdigalian stage of the Early Miocene, roughly 20-15 mya (Baker et al. 2006).

The genera Spheniscus and Eudyptula contain species with a mostly subantarctic distribution centered on South America; some, however, range quite far northwards. They all lack carotenoid coloration, and the former genus has a conspicuous banded head pattern; they are unique among living penguins in nesting in burrows. This group probably radiated eastwards with the Antarctic Circumpolar Current out of the ancestral range of modern penguins throughout the Chattian (Late Oligocene), starting approximately 28 mya (Baker et al. 2006). While the two genera separated during this time, the present-day diversity is the result of a Pliocene radiation, taking place some 4-2 mya (Baker et al. 2006).

The Megadyptes - Eudyptes clade occurs at similar latitudes (though not as far north as the Galapagos Penguin), has its highest diversity in the New Zealand region, and represent a westward dispersal. They are characterized by hairy yellow ornamental head feathers; their bills are at least partly red. These two genera diverged apparently in the Middle Miocene (Langhian, roughly 15-14 mya), but again, the living species of Eudyptes are the product of a later radiation, stretching from about the late Tortonian (Late Miocene, 8 mya) to the end of the Pliocene (Baker et al. 2006).

It is most interesting to note that the geographical and temporal pattern or spheniscine evolution corresponds closely to two episodes of global cooling documented in the paleoclimatic record (Baker et al. 2006). The emergence of the subantarctic lineage at the end of the Bartonian corresponds with the onset of the slow period of cooling that eventually led to the ice ages some 35 million years later. With habitat on the Antarctic coasts declining, by the Priabonian more hospitable conditions for most penguins existed in the subantarctic regions rather than in Antarctica itself. Notably, the cold Antarctic Circumpolar Current also started as a continuous circumpolar flow only around 30 mya, on the one hand forcing the Antarctic cooling, and on the other faciliating the eastward expansion of Spheniscus to South America and eventually beyond (Baker et al. 2006).

Later, an interspersed period of slight warming was ended by the Middle Miocene Climate Transition, a sharp drop in global average temperature from 14 to 12 mya, and similar abrupt cooling events followed at 8 mya and 4 mya; by the end of the Tortonian, the Antarctic ice sheet was already much like today in volume and extent. The emergence of most of today's subantarctic penguin species almost certainly was caused by this sequence of Neogene climate shifts.

Penguin ancestry beyond Waimanu remains unknown and not well resolved by molecular or morphological analyses. The latter tend to be confounded by the strong adative autapomorphies of the Sphenisciformes; a sometimes perceived fairly close relationship between penguins and grebes is almost certainly an error based on both groups' strong diving adaptations, which are homoplasies. On the other hand, different DNA sequence datasets do not agree in detail with each other either.

What seems clear is that penguins belong to a clade of Neoaves (living birds except paleognaths and fowl) which comprises of what is sometimes called "higher waterbirds" to distinguish them from the more ancient waterfowl. This group contains such birds as storks, rails, and the seabirds, with the possible exception of the Charadriiformes (Fain & Houde 2004).

Inside this group, penguin relationships are far less clear. Depending on the analysis and dataset, a close relationship to Ciconiiformes (e.g. Slack et al. 2006) or to Procellariiformes (Baker et al. 2006) has been suggested. Some (e.g. Mayr 2005) think the penguin-like plotopterids (usually considered relatives of anhingas and cormorants) may actually be a sister group of the penguins, and that penguins may have ultimately shared a common ancestor with the Pelecaniformes and consequently would have to be included in that order, or that the plotopterids were not as close to other pelecaniforms as generally assumed, which would necessitate splitting the traditional Pelecaniformes in three

Answer 5

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