is any one know what is the color changes mechanism at the chameleon chromatophore?

Answer 1

Start here, at the Mad Scientist Network (one of my favorite sites), which has links to still more:
http://www.madsci.org/posts/archives/mar97/857252824.Zo.r.html

This explains it in easy-to-understand language:
http://science.howstuffworks.com/animal-camouflage2.htm

There's always Wikipedia:

"Some species can rapidly change colour through mechanisms that translocate pigment and reorient reflective plates within chromatophores. This process, often used as a type of camouflage, is called physiological colour change. Cephalopods such as octopus have complex chromatophore organs controlled by muscles to achieve this, while vertebrates such as chameleons generate a similar effect by cell signaling. Such signals can be hormones or neurotransmitters and may be initiated by changes in mood, temperature, stress or visible changes in local environment." (1)

"Chameleons have specialized cells, collectively called chromatophores, that lie in layers under their transparent outer skin. The cells in the upper layer, called xanthophores and erythrophores, contain yellow and red pigments respectively. Below these is another layer of cells called iridophores (or guanophores), and they contain the colorless crystalline substance guanine. These reflect amongst others the blue part of incident light. If the upper layer of chromatophores appear mainly yellow, the reflected light becomes green (blue plus yellow). A layer of dark melanin containing melanophores is situated even deeper under the reflective iridophores. The melanophores influence the 'lightness' of the reflected light. All these different pigment cells can rapidly relocate their pigments, thereby influencing the color of the chameleon." (2)